Turnovers have a particular way of gnawing at Doug Marrone.
So those in attendance at Ralph Wilson Stadium for Saturday's preseason game against the Tampa Bay Buccaneers may have seen the steam coming out of the Buffalo Bills coach's ears.
The Bills' starting offense turned the ball over twice on its first two possessions. Quarterback EJ Manuel was intercepted on Buffalo's first possession. Manuel was attempting to find Scott Chandler, but the tight end fell down on his route, leading to an easy play for Bucs safety Dashon Goldson.
After a three-and-out for the offense, the Bills lost a fumble on their third possession. Manuel completed a short dump off to running back C.J. Spiller, but as he attempted to make a move, Bucs linebacker Lavonte David punched the ball out. It was recovered by Tampa Bay's Mason Foster.
Marrone spoke earlier in the week about how monitoring the turnover battle would be one of his primary focuses heading into the week.
Special teams struggles, too: Punter Brian Moorman also had a first quarter to forget. Moorman's first punt from the Bills' 9-yard line traveled only 34 yards, giving Tampa Bay possession at the Buffalo 46. Moorman's next punt only netted 33 yards.
Defense holds -- for a time: The Bills' starting defense looked good out of the gate. The Bucs' first three offensive plays all went for losses of yardage. Cornerback Nickell Robey dropped Robert Herron for a loss of 2 yards, linebacker Nigel Bradham dropped running back Doug Martin for a 1-yard loss and Corbin Bryant sacked quarterback Josh McCown on third down. On Tampa Bay's next possession, Bills safety Da'Norris Searcy picked off McCown.