John Miller and Jordan Mills each withstood challenges this summer to remain the Buffalo Bills' starters at right guard and right tackle, respectively.
Coach Sean McDermott said Monday that both would be in the starting lineup for Sunday's regular-season opener against the New York Jets.
Miller, who had been the No. 1 right guard since the Bills made him a third-round draft pick from Louisville in 2015, seemed to be the most in danger of losing his spot after alternating practice and preseason-game reps with free-agent addition Vlad Ducasse. Ducasse was believed to have at least a slight edge by having previously worked with Bills offensive line coach Juan Castillo when they were with the Baltimore Ravens together.
"The competition between Vlad and John has been fun to watch, in terms of watching both of those players put their best foot forward," McDermott said. "Those guys are going to continue to battle."
Mills, who was the Bills' starting right tackle last season, was thought to be in jeopardy of losing his first-string status after the team made Dion Dawkins a second-round draft pick from Temple last April.
"There was good, healthy competition (between Mills and Dawkins) on the field against Detroit (in last Thursday night's preseason-finale)," McDermott said. "So we feel good about our group there, and with the guys also back from last year, with (center) Eric (Wood) and (left guard) Richie (Incognito)."
Left tackle Cordy Glenn, who has battled problems with his left ankle and left foot, started against the Lions and is expected to start Sunday.
"Cordy got some good (playing) time the other night and we need to continue to acclimate him and get him going in terms of his conditioning and all that type of stuff," McDermott said. "He was in over the weekend and did a nice job putting in extra time."
Addressing other lineup matters, McDermott said that E.J. Gaines, whom the Bills acquired as part of the trade that sent wide receiver Sammy Watkins to the Los Angeles Rams, had "the upper hand" for the starting cornerback spot opposite first-round draft pick Tre'Davious White. Kevon Seymour -- who, along with Shareece Wright and Greg Mabin, was thought to have been in the starting-cornerback competition -- was sent to the Carolina Panthers Saturday as part of a deal that allowed the Bills to acquire wide receiver Kaelin Clay.
The coach also said that starting strong-side linebacker Lorenzo Alexander will double as the No. 2 middle linebacker behind Preston Brown.
"We've got all of the confidence in the world in Lorenzo," McDermott said. "He's shown us he can move around in a short amount of time. He's played on the defensive line, he's played on the offensive line, so if anybody can do it, Lorenzo can. It sounds like a cheer, right?"